In Review / Submitted

Novel rabies outbreak in a marine mammal – the Cape fur seal. Gridley T, Barnes A, Blumberg L, et al. (In review; preprint)

Leatherback sea turtles in the Southeast Atlantic Ocean: Multiple threats analysis. Prosdocimi L, Morais M, Tiwari M, et al. (In review)

Using acoustic signatures to understand the endangered Indian Ocean humpback dolphin (Sousa plumbea). Gridley T, Elwen SH, Frainer G, et al. Scientific Reports. (Submitted)

Killer whale genomes suggest admixture and sex-biased genetic dispersal over a broad geographic range (in review). Sarigol, F., A.J. Welch, R. Reisinger, S.H. Elwen, P. Tixier A. Irvine, C. Guinet, G. Lauriano, P.J. N. de Bruyn, A.R. Hoelzel.

Using acoustic signatures to understand the endangered Indian Ocean humpback dolphin (Sousa plumbea). Gridley, T., Elwen, S.H., Frainer, G., Atkins, S., Gennari, E., Vermeulen, E., James, B.S., Dines, S. F. (in review - Scientific Reports).

2026

Killer whales (Orcinus orca) in the Indo-Pacific: a review of their distribution, population structure, and ecology. Lacetera, P., P. Tixier,  I.M. Reeves, , M. Terrapon, A.Barnes, C, Guinet, S.H.Elwen,J. Totterdell, J. Tucker, P. Cross, G. K. Bisset,J. Dickie and J.P.Y. Arnould. .Mammal Review 56 (3), e70046

Genomic Structure and Demographic History of the Brown Fur Seal: From the Ice Age to Current Isolation. Spanjaard CA, Malan A, Gridley T, Arnould JPY, Kirkwood R & Matthee CA. Journal of Biogeography.

Whale killers on the coast of South Africa: killer whale predation on a humpback whale super-group. Seyboth E, Wilkinson C, Elwen SH, Barnes A & Vermeulen E. African Journal of Marine Science.

Recovery of large baleen whales in the South-East Atlantic: Recent increased presence of blue (Balaenoptera musculus) and fin whales (Balaenoptera physalus). James BS, Gridley T, Radford M, Paterson J, Roux J-P & Elwen SH. African Journal of Marine Science.

Low genetic diversity and regional isolation of South Africa's inshore Bryde's whales. Paynee D, Vermeulen E, Penry G, Elwen SH, Matthee C, Andreotti S & Bloomer P. Conservation Genetics.

Unravelling Taxonomic Complexity in Elusive Cetaceans: Mitogenome Insights into Evolutionary History and Cryptic Diversity of Bryde's Whales. Paynee D, Bloomer P, Penry G, et al. Ecology and Evolution.

New Insights: Impacts of Fireworks on Marine Species in Cape Town, South Africa. Probert R, Elwen S, Dines S & Gridley T. The Effects of Noise on Aquatic Life IV.

Assessing the short-term impacts of in-air firework sounds on marine species in Cape Town, South Africa. Probert R, Elwen S & Gridley T. Scientific Reports.
Killer whales (Orcinus orca) in the Indo-Pacific: a review of their distribution, population structure, and ecology.

2025

First confirmed stranding of a blue whale (Balaenoptera musculus) in Africa (Namibia), and its likely death by ship strike. James BS, Gridley T & Elwen SH. Scientific Society of Swakopmund Reports. (In press)

Vulnerability of marine megafauna to global at-sea anthropogenic threats. VanCompernolle M, Elwen SH, Sequeira AMM, et al. Conservation Biology.

Underwater behavior of Southern Hemisphere humpback whales (Megaptera novaeangliae) feeding in super-groups, from video footage. Elwen SH, Hannén R, Benjamin S, Velasquez Rojas C, Probert R & Gridley T. Marine Mammal Science.

An update on the occurrence of humpback whale (Megaptera novaeangliae) super-groups on the west coast of South Africa. Seyboth E, Findlay K, Vogel A, et al. Marine Mammal Science.

Blue Carbon Potential in an eastern boundary Upwelling Zone – A case study in Namibia. Elwen SH, Khan U, Kreiner A, et al. Deep Sea Research II.

Report of Two Rare Cases of Allonursing and Fostering in the Cape Fur Seal (Arctocephalus pusillus pusillus) in Namibia. Martin M, Gridley T, Elwen SH & Charrier I. Marine Mammal Science.

Heaviside's dolphin Cephalorhynchus heavisidii. Abernethy RB & Elwen SH. (Book chapter) In: Coastal Dolphins and Porpoises, 369–394.

2024

Hermie the Humpback Dolphin – A Longitudinal Case Study Characterising the Acoustic Behaviour of an Endangered Indian Ocean Humpback Dolphin (Sousa plumbea). Dines S, Probert R, Gullen A, Elwen SH, Frainer G & Gridley T. JASA Express Letters.

Prevalence and source of entanglements in Cape fur seals (Arctocephalus pusillus pusillus) in South Africa. Ramilo-Henry M, Elwen SH, Barnes A, Ward V & Gridley T. Regional Studies in Marine Science.

Use of acoustic signals in Cape fur seal mother-pup reunions: individual signature, signal propagation and pups' home range. Martin M, Stow J, Gridley T, Immerz A, Elwen SH & Charrier I. Journal of Experimental Biology.

Pygmy Right Whale Caperea marginata (Gray, 1846). Frainer G & Elwen SH. (Book chapter) In: Handbook of the Mammals of Middle and South America. Cetacea and Sirenia.

Southern right whale dolphin Lissodelphis peronii (Gray, 1846). Frainer G. (Book chapter) In: Handbook of the Mammals of Middle and South America. Cetacea and Sirenia.

Hourglass dolphin Sagmatius cruciger (Quoy & Gaimard, 1824). Frainer G & Dellabianca N. (Book chapter, in press) In: Handbook of the Mammals of Middle and South America. Cetacea and Sirenia.

Behavioral repertoire of Lahille's bottlenose dolphins during the "cooperative fishing" with artisanal fishers. Serpa N, Frainer G, dos Santos B, Bortolotto GA & Moreno IB. Behavioral Ecology and Sociobiology. (In review)

Isochrony in barks of Cape fur seal (Arctocephalus pusillus pusillus) pups and adults. Osiecka AN, Fearey J, Ravignani A & Burchardt LS. Ecology and Evolution.

2023

Ocean Sound Essential Ocean Variable Implementation Plan. Tyack PL, Akamatsu T, Boebel O, et al.

Leaping into the future: Current application and future direction of computer vision and artificial intelligence in marine sciences in South Africa. Da Silva C, Samaai T, Kerwath S, et al. Research Ideas and Outcomes.

Vocal cues to assess arousal state of bottlenose dolphins (Tursiops spp.) involved in public presentations. Probert R, James BS, Elwen SH & Gridley T. Journal of Zoological and Botanical Gardens.

Geographical variation in Cape fur seals' in-air vocalizations across Southern Africa (Namibia and South Africa). Martin M, Stow J, Gridley T, Elwen SH & Charrier I. Bioacoustics.

Inter-site variability in the Cape fur seal's behavioural response to boat noise exposure. Martin M, Gridley T, Elwen SH & Charrier I. Marine Pollution Bulletin.

Automatic Detection and Taxonomic Identification of Dolphin Vocalizations using Convolutional Neural Networks for Passive Acoustic Monitoring. Frainer G, Dufourq E, Fearey J, Dines S, Probert R, Elwen SH & Gridley T. Ecological Informatics.

Effect of Environmental Variables on African Penguin Vocal Activity: Implications for Acoustic Censusing. Feist F, Terranova F, Petersen G, et al. Biology.

Evidence of signature whistles produced by Indian Ocean bottlenose dolphins (Tursiops aduncus) in Mozambique. Probert R, Gullan A, Rocha D, Dines S & Gridley T. Bioacoustics.

Cetacean diversity of the eastern South Atlantic Ocean and Vema Seamount detected during a visual and passive acoustic survey, 2019. Elwen S, Fearey J, Ross-Marsh E, Thompson K, Maack T, Webber T & Gridley T. Journal of the Marine Biological Association of the United Kingdom.

Good fences make good neighbours: territorial male Cape fur seals use spatial acoustic map of neighbours. Martin M, Gridley T, Elwen SH & Charrier I. Behaviour.

Diagnosing gastric ulcers in bottlenose dolphins (Tursiops sp.) using gastroscopy and cytology. Buhrmann CA, Gridley T & Oellermann LK. Zoo Biology.

2022

Early onset of postnatal individual vocal recognition in a highly colonial mammal. Martin M, Gridley T, Elwen SH & Charrier I. Proceedings of the Royal Society B: Biological Sciences.

Rostrum abnormalities in the endangered Indian Ocean humpback dolphin (Sousa plumbea) in South Africa. Frainer G, Elwen SH, Dines SF, et al. Integrative Zoology.

Direct observation of killer whales predating on white sharks and evidence of a flight response. Towner A, Kock A, Stopforth C, Hurwitz D & Elwen SH. Ecology. (Watch related video)

Song recordings suggest feeding ground sharing in Southern Hemisphere humpback whales. Schall E, Djokic D, Ross-Marsh E, et al. Scientific Reports.

Methodological considerations for using naturally distinctive vocalisations in mark capture-recapture frameworks. Fearey J, Elwen SH, Distiller G & Gridley T. Marine Mammal Science.

Fear at the top: killer whale predation drives white shark absence at South Africa's largest aggregation site. Towner A, Watson RGA, Kock AA, et al. African Journal of Marine Science.

In-air acoustic repertoire and associated behaviour of wild juvenile crabeater seals during rehabilitation. Shabangu FW, Hofmeyr GJ, Probert R, Connan M, Buhrmann CA & Gridley T. Bioacoustics.

Mutual mother-pup vocal recognition in the highly colonial Cape fur seal: evidence of discrimination of calls with a high acoustic similarity. Martin M, Gridley T, Elwen SH & Charrier I. Animal Cognition.

Temporal patterns in Cape fur seal (Arctocephalus pusillus pusillus) mother and pup attraction calls. Osiecka AN, Gridley T & Fearey J. Belgian Journal of Zoology.

Citizen science noninvasive genetics reveals population changes in a whale breeding ground. Neveceralova P, Carroll E, Steel D, et al. Global Ecology and Conservation.

Assessment of the impact of anthropogenic airborne noise on the behaviour of Cape fur seals during the breeding season in Namibia. Martin M, Gridley T, Elwen SH & Charrier I. Journal of Experimental Marine Biology and Ecology.

Dorsal fin mark changes for assigning sexes to individual bottlenose dolphins, Tursiops truncatus. James BS, Gridley T, McGovern B, Fearey JL & Elwen SH. Marine Mammal Science.

Detection of humpback whale (Megaptera novaeangliae) non-song vocalisations around the Vema Seamount, Southeast Atlantic. Ross-Marsh EC, Elwen SH, Fearey J, Thompson K, Maack T, Webber T & Gridley T. JASA.

Feel the beat: Cape fur seal males encode their arousal state in their bark rate. Martin M, Gridley T, Elwen SH & Charrier I. The Science of Nature.

HydroMoth: testing a prototype recorder for low-cost passive acoustic monitoring in aquatic ecosystems. Gordon T, Chapuis L, William B, et al. Remote Sensing in Ecology and Conservation.

Sounding out a continent: Seven decades of bioacoustics research in Africa. Becker F, Shabangu F, Gridley T, Wittmer H & Marsland S. Bioacoustics.

2021

Micro-geographic variations and intra-species acoustic partitioning of the vocal repertoire of an extremely colonial pinniped species, the Cape fur seal. Martin M, Gridley T, Elwen SH & Charrier I. Royal Society Open Science. 8. 202241.

Using Signature Whistles to Investigate Population Dynamics of a Threatened Population of Bottlenose Dolphins (Tursiops truncatus). Fearey JL, Elwen SH, Dines S, James BS & Gridley T. Journal of the Namibian Scientific Society.

Vocal correlates of arousal in bottlenose dolphins (Tursiops spp.) in human care. Probert R, Bastian A, Elwen SH, James BS & Gridley T. PLOS ONE. https://doi.org/10.1371/journal.pone.0250913

Predatory species left stranded following the collapse of the sardine Sardinops sagax (Pappe, 1854) stock off the northern Benguela Ecosystem: A review. Erasmus VN, Currie B, Iitembu J, Roux J-P, Elwen SH, Moses K & Tjizoo B. Journal of Marine Systems. Volume 224, December 2021, 103623

South African marine citizen science – benefits, challenges and future directions. Potts W.M., J.B. Mann-Lang, B.Q. Mann, C.L. Griffiths, C.G. Attwood, A.D. de Blocq, S.H. Elwen, R. Nel, K. Sink and R Thornycroft. (2021). African Journal of Marine Science. Volume 43,

Entanglement of Cape fur seals (Arctocephalus pusillus pusillus) at colonies in central Namibia. 2021. Curtis S, Elwen SH, Dreyer NN & Gridley T. Marine Pollution Bulletin. V171. 112759

Science alone won't do it! South Africa's endangered humpback dolphins Sousa plumbea face complex conservation challenges. Plön S, et al. (13 co-authors) Frontiers in Marine Science.

Trophic interactions and feeding ecology of two delphinids in the Benguela upwelling ecosystem inferred from stable isotopes. Caputo M, Elwen SH, Gridley T, Kohler SA, Roux J-P, Froneman PW & Kiszka JJ. Marine Ecology Progress Series.

Extreme ecological constraints lead to high degree of individual stereotypy in the vocal repertoire of the Cape fur seal (Arctocephalus pusillus pusillus). Martin M, Gridley T, Elwen SH & Charrier I. Behavioral Ecology and Sociobiology, 75, 1–16.

Natal origin of Namibian gray whale reveals new distance record for in-water migration. Hoelzel AR, Sarigol F, Gridley T & Elwen SH. Biology Letters.

Echolocation click parameters and biosonar behaviour of the dwarf sperm whale (Kogia sima). Malinka CE, Tønnesen P, Dunn CA, Claridge DE, Gridley T, Elwen SH & Madsen PT. Journal of Experimental Biology.

2020

Towed passive acoustic monitoring complements visual survey methods for Heaviside's dolphins Cephalorhynchus heavisidii in the Namibian Islands Marine Protected Area. Gridley T, Martin MJ, Swift RJ, Roux J-P, Slater J & Elwen SH. African Journal of Marine Science.

First abundance estimates of Heaviside's (Cephalorhynchus heavisidii) and dusky (Lagenorhynchus obscurus) dolphins off Namibia using a novel visual and acoustic line transect survey. Martin MJ, Gridley T, Roux J-P & Elwen SH. Frontiers in Marine Science.

A first look at humpback whale (Megaptera novaeangliae) song structure from western South Africa. Hawkey JS, Elwen SH, James BS, Prinsloo AS & Gridley T. African Zoology. 55 (3)

Mark-recapture of individually distinctive acoustic labels – a case study with bottlenose dolphins (Tursiops truncatus). Longden E, Elwen SH, McGovern B, James BS, Embling C & Gridley T. Journal of Mammalogy.

Prolonged nursing in Cape fur seals (Arctocephalus pusillus pusillus) at Cape Cross colony, Namibia. 2020. Osiecka A, Fearey J, Elwen SH & Gridley T. African Zoology. 55(3). https://doi.org/10.1080/15627020.2020.1768144

Singing in South Africa: Monitoring the occurrence of humpback whale song near the Western Cape. Ross-Marsh EC, Elwen SH, Prinsloo AS, James BS & Gridley T. Bioacoustics.

2019

Risky Business: repeated live strandings of common bottlenose dolphins (Tursiops truncatus) while foraging in a shallow water environment. 2019. McGovern B, Gridley T, James BS & Elwen SH. Marine Mammal Science. DOI: 10.1111/mms.12634

A synthesis of three decades of socio-ecological change in False Bay, South Africa: setting the scene for multidisciplinary research and management. 2019 Pfaff M, Turpie J, Krug M, et al. Elementa: Science of the Anthropocene. Elem Sci Anth, 7: 32. DOI: https://doi.org/10.1525/elementa.367  ALTMETRIC Score currently 47

Identification of potential signature whistles from free-ranging common dolphins (Delphinus spp.), in South Africa. 2021 Fearey J, Elwen SH, James BS & Gridley T. Animal Cognition. https://doi.org/10.1007/s10071-019-01274-1

Predicting habitat suitability for cetaceans in Namibia using MinxEnt. 2019 De Rock P, Elwen SH, Roux J-P, Leeney RH, James BS, Visser V, Martin MJ & Gridley T. Marine Ecology Progress Series. doi: 10.3354/meps12934

To buzz or burst-pulse? The functional role of Heaviside's dolphin (Cephalorhynchus heavisidii) rapidly pulsed signals. 2019. Martin MJ, Elwen SH, Kassanjee R & Gridley T. Animal Behaviour. https://doi.org/10.1016/j.anbehav.2019.01.007

Abundance estimates of an isolated population of common bottlenose dolphins in Walvis Bay, Namibia 2008-2012. 2019. Elwen SH, Leeney RH & Gridley T. African Journal of Marine Science 41(1). 61-70 https://doi.org/10.2989/1814232X.2019.1572538

2018

Hybridisation in bottlenose dolphins – A case study of Tursiops aduncus × T. truncatus hybrids and successful back-cross hybridisation events Gridley T, Elwen SH, Harris G, Moore DM, Hoelzel AR & Lampen F. (2018). PLOS ONE, 13(9), e0201722.

Heaviside's dolphins (Cephalorhynchus heavisidii) relax acoustic crypsis to increase communication range Martin MJ, Gridley T, Elwen SH & Jensen FH. (2018). Proceedings of the Royal Society B, 285(1883), 20181178.

The host-specific whale louse (Cyamus boopis) as a tool for interpreting humpback whale (Megaptera novaeangliae) migratory routes Iwasa-Arai T, Serejo C, Siciliano S, Ott PH, Freire AS, Elwen SH, Crespo EA, Colosio AC, Carvalho VL & Rodríguez-Rey GT. (2018). Journal of Experimental Marine Biology and Ecology, 505, 45–51.

Song recorded from a super-group of humpback whales on a mid-latitude feeding ground Gridley T, Pereira Silva MF, Wilkinson C, Seakamela SM & Elwen SH. (2018). JASA Express Letters, 143(4), EL298–EL304.

2017

Indian Ocean humpback dolphin (Sousa plumbea) movement patterns along the South African coast Vermeulen E, Bouveroux T, et al. (14 authors). (2017). Aquatic Conservation: Marine and Freshwater Ecosystems, 28(1), 231–240. Automatic classification of whistles from coastal dolphins of the southern African subregion Erbs F, Elwen SH & Gridley T. (2017). Journal of the Acoustical Society of America, 141(4), 2489–2500.

2016

Impacts of acoustic identity pinger tags on bottlenose dolphins (Tursiops truncatus) Elwen SH, McGovern B, Tregenza N & Gridley T. (2016). Proceedings of Meetings on Acoustics, 27(1), 010040. Bottlenose dolphins change their whistling characteristics in relation to vessel presence, surface behavior and group composition Gridley T, Elwen SH, Rashley G, Badenas-Krakauer A & Heiler J. (2016). Proceedings of Meetings on Acoustics, 27(1), 010030. Changes in bottlenose dolphin whistle parameters related to vessel presence, surface behaviour and group composition Heiler J, Elwen SH, Kriesell HJ & Gridley T. (2016). Animal Behaviour, 117, 167–177.

2015

Epidemiology of lobomycosis-like disease in bottlenose dolphins from South America and southern Africa Van Bressem M-F, Simões-Lopes PC, Félix F, Kiszka JJ, Daura-Jorge FG, Avila IC, Secchi ER, Flach L, Fruet PF, du Toit K, Ott PH, Elwen S, Di Giacomo AB, Wagner J, Banks A & Van Waerebeek K. (2015). Diseases of Aquatic Organisms, 117(1), 59–75.

The acoustic repertoire of wild common bottlenose dolphins (Tursiops truncatus) in Walvis Bay, Namibia Gridley T, Kriesell H, Nastasi A & Elwen SH. (2015). Bioacoustics, 24(2), 153–174.

Abundance and degree of residency of humpback dolphins (Sousa plumbea) in Mossel Bay, South Africa James BS, Bester MN, Penry GS, Gennari E & Elwen SH. (2015). African Journal of Marine Science, 37(3), 383–394.

The use of data from a Platform of Opportunity (Whale Watching) to study coastal cetaceans on the south west coast of South Africa Vinding KV, Bester MN, Kirkman SP, Chivell W & Elwen SH. (2015). Tourism in Marine Environments, 11(1), 33–54.

Possible non-offspring nursing in the southern right whale, Eubalaena australis Best PB, Elwen SH, Palsbøll PJ, Thornton M, Austin E & Vinding K. (2015). Journal of Mammalogy, 96(2), 405–416.

2014

Humpback whales off Namibia: occurrence, seasonality and a regional comparison of photographic catalogues and scarring Elwen SH, Tonachella N, Barendse J, Collins T, Best PB, Rosenbaum HC, Leeney RH & Gridley T. (2014). Journal of Mammalogy, 95(5), 1064–1076.

Identification of signature whistles in wild bottlenose dolphins (Tursiops truncatus) from Namibia Kriesell HJ, Elwen SH, Nastasi A & Gridley T. (2014). PLOS ONE, 9(9), e106317.

2013

False killer whale (Pseudorca crassidens) sightings in continental shelf habitat off Gabon and Côte d'Ivoire (Africa) Weir CR, Collins T, Cross T, Gill A, Elwen SH, Unwin M & Parnell RJ. (2013). Marine Biodiversity Records, 6, e65.

Pygmy right whale records from Namibia Leeney RH, Post K, Best PB, Hazevoet CJ & Elwen SH. (2013). African Journal of Marine Science, 35(1), 133–139.

Records of Kogiid whales in Namibia including the first record of the dwarf sperm whale (K. sima) Elwen SH, Gridley T, Roux J-P, Best PB & Smale MJ. (2013). Marine Biodiversity Records, 6, e45.

Integrating multiple data sources to assess the distribution and abundance of bottlenose dolphins (Tursiops truncatus) in Scottish waters Cheney B, Thompson PM, Ingram SN, Hammond PS, Stevick PT, Durban JW, Culloch RM, Elwen SH, Mandleberg L, Janik VM, Quick NJ, Islas-Villanueva V, Robinson KP, Costa M, Eisfeld SM, Walters A, Phillips C, Weir CR, Evans PGH, Anderwald P, Reid RJ, Reid JB & Wilson B. (2013). Mammal Review, 43(1), 71–88.

2012

Death under the surface – Cape fur seal predating on a southern giant petrel in Walvis Bay, Namibia Elwen SH, Meintjies J & Roux J-P. (2012). Ornithological Observations, 3, 206–210.

2011

The first African Marine Mammal Colloquium, South Africa, May 2010 Elwen SH, Thornton M, Kirkman SP, Pistorius P & Weir CR. (2011). African Journal of Marine Science, 33(3), 349–351.

Cetacean research in the southern African subregion: a review of previous studies and current knowledge Elwen SH, Findlay KP, Kiszka J & Weir CR. (2011). African Journal of Marine Science, 33(3), 469–493.

Transit station or destination? Attendance patterns, regional movement, and population estimate of humpback whales Megaptera novaeangliae off West South Africa based on photographic and genotypic matching Barendse J, Best PB, Thornton M, Elwen SH, Rosenbaum HC, Carvalho I, Pomilla C, Collins TJQ & Meÿer MA. (2011). African Journal of Marine Science, 33(3), 353–373.

Distribution and abundance of humpback whales, Megaptera novaeangliae, off the coast of Mozambique, 2003 Findlay KP, Meÿer M, Elwen SH, Kotze D, Johnson R, Truter P, Uamusse C, Sitoe S, Wilke C, Kerwath S, Swanson S, Staverees L & van der Westhuizen J. (2011). Journal of Cetacean Research and Management, Special Issue 3, 163–174.

Interactions between leatherback turtles and killer whales in Namibian waters, including predation Elwen SH & Leeney RH. (2011). South African Journal of Wildlife Research, 41(2), 205–209.

Using Static Acoustic Monitoring to describe Echolocation Behaviour of Heaviside's Dolphins in Namibia Leeney RH, Carslake D & Elwen SH. (2011). Aquatic Mammals, 37(2), 151–160.

2010 and earlier

Injury and Subsequent Healing of a Propeller Strike Injury to a Heaviside's dolphin (Cephalorhynchus heavisidii) Elwen SH & Leeney RH. (2010). Aquatic Mammals, 36(4), 382–387.

Near-shore distribution of Heaviside's (Cephalorhynchus heavisidii) and dusky dolphins (Lagenorhynchus obscurus) at the southern limit of their range in South Africa Elwen SH, Best PB, Thornton M & Reeb D. (2010). African Zoology, 45(1), 78–91.

Diurnal movements and behaviour of Heaviside's dolphins (Cephalorhynchus heavisidii), with some comparative data for dusky dolphins (Lagenorhynchus obscurus) Elwen SH, Best PB, Reeb D & Thornton M. (2009). South African Journal of Wildlife Research, 39(2), 143–154.

A population estimate of Heaviside's dolphins Cephalorhynchus heavisidii in the southern end of their range Elwen SH, Reeb D, Thornton M & Best P. (2009). Marine Mammal Science, 25(1), 107–124.

Range and movements of a nearshore delphinid, Heaviside's dolphin Cephalorhynchus heavisidii as determined from satellite telemetry Elwen SH, Meÿer MAM, Best PB, Kotze PGH, Thornton M & Swanson S. (2006). Journal of Mammalogy, 87(5), 866–877.

Environmental factors influencing the distribution of southern right whales (Eubalaena australis) on the south coast of South Africa I: Broad scale patterns Elwen SH & Best PB. (2004). Marine Mammal Science, 20(3), 567–582.

Are there reproductive benefits to southern right whale females (Eubalaena australis) associated with their coastal distribution off South Africa? Elwen SH & Best PB. (2004). Marine Ecology Progress Series, 269, 289–295.

Book Chapters & Reviews

Book Review: Ethology and Behavioural Ecology (Editor B. Würsig). Gridley T & Elwen SH. (2019) Marine Mammal Science.
Patterns of sympatry in Lagenorhynchus and Cephalorhynchus dolphins in different habitats. Heinrich S, Elwen SH & Brager S. (2010) In: Dusky Dolphins: Master Acrobats on Different Shores (Eds. Würsig B & Würsig M.), pp 313–331. Academic/Elsevier Press.